9-segmented Cornuproetinae of Morocco
Order Proetida
Family Proetidae
Subfamily Cornuproetinae
A number of cornuproetine genera is characterized by having 9 instead of 10 thorax segments. Whereas Sculptoproetus and Lepidoproetus were described by Erben more than half a centurary ago, Timsaloproetus was recently recognised by Gibb & Chatterton. We added another new genus, Hollandiella, which was first discovered by the late Dieter Holland (hence its name). Many of these species are very small in size (with a notable exception of some species of Lepidoproetus).
Family Proetidae
Subfamily Cornuproetinae
A number of cornuproetine genera is characterized by having 9 instead of 10 thorax segments. Whereas Sculptoproetus and Lepidoproetus were described by Erben more than half a centurary ago, Timsaloproetus was recently recognised by Gibb & Chatterton. We added another new genus, Hollandiella, which was first discovered by the late Dieter Holland (hence its name). Many of these species are very small in size (with a notable exception of some species of Lepidoproetus).
Sculptoproetus
Name: Sculptoproetus raki Van Viersen & Lerouge, 2019
Stratigraphy: 'Cyphaspides couche', late Emsian
Locality: Bizmah (Rissani), Morocco
Very small Sculptoproetus with notably large eyes and a fine fingerprint-like sculpture of terrace lines with granules. This species was named after Štěpán Rak from the Museum of the Czech Karst. Notice the similarities to Sculptoproetus africanus, formerly Macroblepharum.
Stratigraphy: 'Cyphaspides couche', late Emsian
Locality: Bizmah (Rissani), Morocco
Very small Sculptoproetus with notably large eyes and a fine fingerprint-like sculpture of terrace lines with granules. This species was named after Štěpán Rak from the Museum of the Czech Karst. Notice the similarities to Sculptoproetus africanus, formerly Macroblepharum.
Timsaloproetus
Name: Timsaloproetus pulchistriatus repens van Viersen & Lerouge, 2019
Stratigraphy: Timrhanrhart Formation, upper Emsian
Locality: Foum Zguid, Morocco
Stratigraphy: Timrhanrhart Formation, upper Emsian
Locality: Foum Zguid, Morocco
|
|
Lepidoproetus
Name: Lepidoproetus maharchianus (Johnson & Fortey, 2012)
Stratigraphy: Ihandar formation, Pragian
Locality: Maharch
A spineless Lepidoproetus, showing a very fine fingerprint-like sculpture on the carapace. Broad, medially protruding anterior border and tongue-shaped glabella.
Stratigraphy: Ihandar formation, Pragian
Locality: Maharch
A spineless Lepidoproetus, showing a very fine fingerprint-like sculpture on the carapace. Broad, medially protruding anterior border and tongue-shaped glabella.
|
|
Holandiella
Name: Hollandiella lebruni van Viersen & Lerouge, 2019
Stratigraphy: Timrhanrhart Formation, Emsian part
Locality: Lansser, Morocco
This species was named after Patrice Lebrun, in recognition of his work on Moroccan trilobites.
Stratigraphy: Timrhanrhart Formation, Emsian part
Locality: Lansser, Morocco
This species was named after Patrice Lebrun, in recognition of his work on Moroccan trilobites.
Name: Hollandiella verecunda van Viersen & Lerouge, 2019
Stratigraphy: 'Lobopyge couche', Pragian
Locality: Tafraoute, Morocco
Stratigraphy: 'Lobopyge couche', Pragian
Locality: Tafraoute, Morocco
Name: Hollandiella curvirostris van Viersen & Lerouge, 2019
Stratigraphy: Ihandar formation, Pragian
Locality: Oued Atchana, Morocco
This is the holotype of the type species of this genus. Hollandiella was recently described as a new cornuproetine genus with 9 thorax segments, in honour of the late Dieter Holland.
Stratigraphy: Ihandar formation, Pragian
Locality: Oued Atchana, Morocco
This is the holotype of the type species of this genus. Hollandiella was recently described as a new cornuproetine genus with 9 thorax segments, in honour of the late Dieter Holland.
References
Chatterton B., Fortey R., Brett K., Gibb S., McKellar R. 2006. Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jbel Gara el Zguilma, southern Morocco. Paleontographica Canadiana 25, 182pp.
Gibb, S.L., and B.D.E. Chatterton. 2007. Timsaloproetus new genus (Proetida: Trilobita) and included species from Lower and Mid- dle strata of southern Morocco. Journal of Paleontology 81: 352–367.
Johnson, R.G., and R.A. Fortey. 2012. Proetid trilobites from the Lower Devonian (Pragian) Ihandar Formation, Anti-Atlas, Morocco. Journal of Paleontology 86: 1032–1050.
Viersen, A.P. van & Lerouge, F., 2019. Cornuproetine (proetide) trilobites with nine thorax segments from the Devonian of Morocco, Germany and the Czech Republic. PalZ (Paläontologische Zeitschrift) 94 (2): 227-254. Published online: 26 October 2019.
Viersen, A.P. van & Lerouge, F. 2021. Timsaloproetus alissae sp. nov. (Trilobita: Proetidae) from the Lower Devonian of southern Morocco. PalZ (Paläontologische Zeitschrift). https://doi.org/10.1007/s12542-020-00543-6. [ full-text ]
Chatterton B., Fortey R., Brett K., Gibb S., McKellar R. 2006. Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jbel Gara el Zguilma, southern Morocco. Paleontographica Canadiana 25, 182pp.
Gibb, S.L., and B.D.E. Chatterton. 2007. Timsaloproetus new genus (Proetida: Trilobita) and included species from Lower and Mid- dle strata of southern Morocco. Journal of Paleontology 81: 352–367.
Johnson, R.G., and R.A. Fortey. 2012. Proetid trilobites from the Lower Devonian (Pragian) Ihandar Formation, Anti-Atlas, Morocco. Journal of Paleontology 86: 1032–1050.
Viersen, A.P. van & Lerouge, F., 2019. Cornuproetine (proetide) trilobites with nine thorax segments from the Devonian of Morocco, Germany and the Czech Republic. PalZ (Paläontologische Zeitschrift) 94 (2): 227-254. Published online: 26 October 2019.
Viersen, A.P. van & Lerouge, F. 2021. Timsaloproetus alissae sp. nov. (Trilobita: Proetidae) from the Lower Devonian of southern Morocco. PalZ (Paläontologische Zeitschrift). https://doi.org/10.1007/s12542-020-00543-6. [ full-text ]